The peer-reviewed quarterly journal Systematics and Biodiversity recently published the report – Perspectives. Colour and size variation in Junonia villida ( Lepidoptera, Nymphalidae ) : subspecies or phenotypic plasticity ? by R.I. Vane-Wright and W. John Tennant.
The review was based on” examination of c.1500 museum specimens from its entire geographical range from the Chagos Islands in the Indian Ocean to the Gambiers in mid-Pacific, spanning over 154 degrees of longitude, or 43% of the circumference of the Earth. Mostly found in open grasslands or disturbed areas, including woodlands and disturbed forests from sea-level up to 1500m or more in elevation, it is variable in wing colour pattern on both upper and lower surfaces. The causes of this variability are uncertain, but temperature, photoperiod, rainfall, migration and perhaps underlying geographical differentiation may all play a role.”
The Clarence Valley Meadow Argus butterfly is represented by the image on lower right side taken on the Conservation Area for the cover illustration showing six Australian butterfly with various differences in colour pattern.
Further reference available; http://www.tandfonline.com/toc/tsab20/current